Research Interests:

• Demography
• Population Dynamics
• Population Genetics
• Optimality modelling
• Philosophy of biology
• Conservation Biology

Education:

• B.A. in Biology, The University of Rochester, 1978
• Ph.D. in Biology, Harvard University,1984

Grants:

• National Institutes of Health Postdoctoral Fellowship, 1986 - 1989
• National Science Foundation Grant, Molecular Evolution and
• Sex Ratio Evolution in the genus Nasonia, 1996 - 2000
• National Science Foundation Grant, Adaptation, Phylogenetic Inertia, and Trait Independence, 2002 - 2003
• National Science Foundation, Algorithmic Problems in Population-Scale Genomics, 2002 - 2005
• National Institute of Aging Program Project Grant, Biodemographic Determinants of Life Span, 2003 - 2008
• National Institute of Health Grant, Genetic Studies of Substance Abuse in Iowa Adoptees, 2004 - 2009
• National Science Foundation Grant, Computational Population Genomics: Using Variation to Connect Genotypes to Phenotypes, 2005 - 2008

Publications

Senior Research Scientist

Research Interests:

• Aculeate Hymenoptera
• Insect Behavior
• Sex Allocation
• Systematics
• Phylogeny

Education:

• B.A., University of California at Riverside, 1969
• M.A., University of California at Riverside, 1973
• Ph.D., Department of Biology, Harvard University, 1980

Publications